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White-capped Albatross Thalassarche cauta Scientific name definitions

Josep del Hoyo, Carles Carboneras, Francesc Jutglar, Nigel Collar, and Guy M. Kirwan
Version: 1.0 — Published March 4, 2020
Text last updated January 3, 2017

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Introduction

Taxonomic note: Lump. This account is a combination of multiple species accounts originally published in HBW Alive. That content has been combined and labeled here at the subspecies level. Moving forward we will create a more unified account for this parent taxon. Please consider contributing your expertise to update this account.

Field Identification

White-capped Albatross (cauta)

90–100 cm (1); male 3900–5100 g, female 3200–4400 g (2); 210–260 cm (1). Largest and stoutest-billed of mollymawks , with most extensive white underwing and diagnostic squarish black patch where leading edge meets body. Adult has white cap contrasting with grey upper face  and especially with blackish ‘eyebrow’ , narrow white crescent framing rear of eye, the pale grey tinge often extending diffusely over lower face and hindneck to uppermost mantle, becoming dark grey on lower mantle and back, blackish grey on scapulars and upperwing (dark upperparts browner in worn plumage), outer primaries with white shafts, rump and uppertail-coverts white; tail dark grey; underwing clean white with diagnostic small blackish patch at very base of leading edge and especially narrow dark fringe on both leading and trailing edges, slightly broader at carpal bend, narrow on wingtip because of broadly white bases to primaries, can be virtually absent at proximal base of leading edge; underparts white; iris dark brown; bill grey (can have slight greenish or olive tinge), with yellow tip  and often yellow base to culmen, culminicorn usually has narrow black frame separating it from latericorn and forehead, often visible only at base, narrow and inconspicuous yellow along cutting edges, more noticeable narrow orange line at very base of mandible; pink gape line (usually concealed); legs pale pinkish with some grey tones. Sexes similar, but female is marginally smaller in most measurements (2); > 80% of adults can be sexed based on wing chord, maximum head length and bill measurements (3). Albinos have been recorded  . Juvenile very similar but distinguished by grey bill with black tip , darker and more brownish tinge on neck, especially at rear and on sides, where may form collar, sometimes contrasting with paler head, but little or no contrast between cap and face. Separated from similar species  by head and bill colour; <em>T. salvini</em> has less white crown, thus appears less capped, although forecrown still contrasts with foreface, and pale grey extends through face and foreneck giving it more hooded appearance, bill greyish to horn , with culmen and tip of maxilla cream to straw-yellow, mandible with blackish tip, the lower ridge tends to be cream-coloured and with narrow orange line at very base. T. eremita is strongly hooded, with darker grey of head  and neck  extending over entire cap, usually paler on central foreneck, bill strong but shorter, brighter than that of salvini and present species, yellow with brownish to olive tinge, culmen purer straw-yellow to rich cream-yellow, sometimes rather yolk-yellow overall, the mandible with black tip and narrow orange line at base; both salvini and present species have darker undersurface to primaries, thus dark margin to underwing looks broader at wingtip.

White-capped Albatross (steadi)

90–100 cm (1); 2600–5300 g (1). Mid-sized black, slate-grey and white albatross with black thumbmark at base of leading edge of underwing  . Bold white cap contrasting with pale silver-grey  face  and darker brow  . Some adults can have very white back with brown tips to feathers; mantle  dark grey and tail black; body all white; bill  pale grey/blue with yellow tip  . Sexes alike, but male averages larger in most measurements (2, 1). Juvenile  has grey bill with dark tip, which lightens with age, and darker head than adult, with grey extending to collar. Very similar to slightly smaller <em>T. cauta</em> , but present species has paler face  and less yellow on culmen. <em>T. salvini</em> and, especially, <em>T. eremita</em> have darker grey heads  and lack white cap.

Systematics History

Editor's Note: This article requires further editing work to merge existing content into the appropriate Subspecies sections. Please bear with us while this update takes place.

White-capped Albatross (cauta)

Previously considered conspecific with T. steadi, T. eremita and T. salvini (but see those species); in past, T. steadi was often considered doubtfully valid even as a subspecies. Monotypic.

White-capped Albatross (steadi)

Until recently considered conspecific with T. cauta, T. eremita and T. salvini (see latter two below); in past, was often treated as a synonym of cauta, and normally still considered conspecific (4, 5); indeed, differences from cauta are small (genetic distance only 0·2% (6) ), involving greater size (on basis of published evidence (7), effect size for wing 2.12, score 2); no yellow at base of bill of adult vs typically yellow bill base (7, 8) ) (2); paler grey cheek (1); slightly later breeding season (Nov–Dec vs Sept–Oct) (no score accepted here). A score of 5 suggests subspecies status, but taxon accepted by ACAP (Agreement on the Conservation of Albatrosses and Petrels) as a full species, largely on account of earlier genetic studies (9, 10); pending further clarifications, this arrangement provisionally and precautionarily accepted here. Monotypic.

Subspecies


EBIRD GROUP (MONOTYPIC)

White-capped Albatross (cauta) Thalassarche cauta cauta Scientific name definitions

Distribution

S Australasian waters, breeding on three small islands off Tasmania (Albatross, Mewstone and Pedra Branca) (11).

EBIRD GROUP (MONOTYPIC)

White-capped Albatross (steadi) Thalassarche cauta steadi Scientific name definitions

Distribution

S Atlantic and S Indian Oceans to Australasian waters, breeding in Auckland and Antipodes Is.

Distribution

White-capped Albatross (cauta)

S Australasian waters, breeding on three small islands off Tasmania (Albatross, Mewstone and Pedra Branca) (11).

White-capped Albatross (steadi)

S Atlantic and S Indian Oceans to Australasian waters, breeding in Auckland and Antipodes Is.

Habitat

White-capped Albatross (cauta)

Marine ; less pelagic than many albatrosses, frequently occurring over continental shelf and even close inshore. Breeds on offshore islands occupying slopes or flatter ground, in areas with boulders but generally sparse vegetation.

White-capped Albatross (steadi)

Marine ; less pelagic than many albatrosses, frequently occurring over continental shelf and even close inshore. Breeding colonies  generally located on rock islands, in sheltered crevices and well-vegetated slopes  .

Migration Overview

White-capped Albatross (cauta)

Clear segregation between birds from different Tasmanian colonies during breeding season, thus breeders on Albatross I, at W end of Bass Strait, forage over Australian continental shelf or slope waters off NW Tasmania, while those nesting on Pedra Branca, off S Tasmania, feed E of Tasmania, between the colony and SE edge of continental shelf, generally within 265 km of the colonies (12). Post-breeding adults also generally remain comparatively close to breeding areas, over continental shelf and slope off SE Australia and generally continue to visit colony through much of the non-breeding period (13), whereas young birds, in first five years of life, are considerably more mobile, with those from Mewstone colony, close to Pedra Branca, moving as far as South Africa and Namibia (14), where concentrates off S & W coasts, and birds from Albatross I have been recovered off Western Australia (2). Uncommon at sea around Macquarie I between austral summer and autumn (15). Either this species or T. steadi is commonly observed off Amsterdam and St Paul in boreal winter (1, 16), with single records from Zanzibar (17) and Mauritius (the latter could also have been T. salvini) (18), and ‘shy-type’ albatrosses have also been recorded in reasonable numbers in S & SW Atlantic, off Tristan da Cunha (19), South Georgia (20), Falklands (21), Argentina (22), Uruguay (23) and S Brazil, exceptionally N to Bahia (24); most of these records apparently involve steadi, but there are some claims of the present species, although doubts persist (25). Status in New Zealand waters also unclear due to extreme difficulty of separating this species from T. steadi (26). One banded bird from Albatross I recovered in N New Zealand. Records in N Hemisphere off Pacific coast of North America (California, Aug–Sept 1999; Washington, Jan 2000; Oregon, Oct 2001) (27), as well as in N Red Sea (Eilat, Israel, Feb–Mar 1981) and off Somalia (11º N, Sept 1986) (28).

White-capped Albatross (steadi)

Tracking studies, bird band recoveries and DNA-based identification of bycatch specimens have confirmed that the species forages in Tasmanian water and South African  /Namibian waters, and immatures are now thought to occur regularly throughout the S Atlantic and SW Indian Oceans. Thus, either this species or T. cauta is commonly observed off Amsterdam and St Paul in boreal winter (1, 16), with single records from Zanzibar (17) and Mauritius (the latter could also have been T. salvini) (18), and ‘shy-type’ albatrosses have also been recorded in reasonable numbers in S & SW Atlantic, off Tristan da Cunha (19), South Georgia (20, 29), Falklands (21), Argentina (22) (in which country there is one definite—specimen—record of steadi) (30), Uruguay (23) and S Brazil (where steadi also recently confirmed from a specimen) (31), exceptionally N to Bahia (24); as mentioned, most of records of ‘shy-type’ albatrosses do apparently involve steadi (including a bird attending a colony of T. melanophris on South Georgia in Feb 2003 and 2004) (29), but there are some claims of T. cauta, although doubts generally persist concerning these (25). One N Hemisphere record, an adult collected off Washington state in Sept 1951 (27).

Diet and Foraging

White-capped Albatross (cauta)

Mostly cephalopods (Nototodarus sloani, Sepioteuthis australis) and fish (Sardinops neopilchardus, Engraulis australis, Trachurus declivis); also fish offal, barnacles and other crustaceans; study of chick diet on Albatross I (Tasmania) found that fish (especially Trachurus declivis and Emmelichthys nitidus) dominated by weight (89%), with cepahalopods comprising most of the remainder (10%) (2). Obtains food by surface-seizing or diving, often from air in shallow plunge (latter usually < 5 seconds), and most dives (87%) are within 3·5 m of surface, but reaching maximum of 7·4 m (32). Has been recorded taking Trachurus declivis directly in flight low over water (2). Some feeding occurs at night. Often feeds in company of other Procellariiformes, and sometimes follows cetaceans. Despite its name, attends fishing boats, where competes greedily for food.

White-capped Albatross (steadi)

Dietary information much less detailed than for T. cauta. Main foods include, fish  , cephalopods, crustaceans and tunicates. Generally feeds by surface-seizing, but may undertake shallow surface-dives. Regularly follows ships and fish-processing discharge  (33) apparently comprises a significant proportion of the species’ diet. Has been observed associating with other seabirds and dusky dolphins (Lagenorhynchus obscurus) (34).

Sounds and Vocal Behavior

White-capped Albatross (cauta)

Typical array of croaks, groans and wails (2). Differences between this species and other members of the ‘shy albatross’ group, if any, have yet to be clearly established and require further research (2).

White-capped Albatross (steadi)

No known differences in vocalizations  between present species and T. cauta (1).

Breeding

White-capped Albatross (cauta)

Annual in most instances, starting Sept, with egg-laying mostly in second half of this month, hatching mostly first half of Dec and fledging and colony desertion in late Apr (2). No data concerning permanence of pair-bond (1). Forms colonies of variable density; nest  is large mound of mud, guano, feathers, grass and roots (2). Single white egg with reddish-brown spots at large end (1), size 95–121 mm × 60–94 mm (2); incubation  68–75 days with stints of 1–7 days (but rarely > 5 days) (2); chicks  have white down, brooded and guarded c. 3–4 weeks (2), thereafter fed comparatively small meals of 372 g, every 22 hours at equal rates by males and females (though latter undertake longer foraging trips) (13), with mean meal size increasing but feeding rate decreasing with time, and receiving an estimated 44·9 kg of food over entire period (35); fledging c. 112–136 (mean 127) days at 5–5·7 kg; male chicks are fed c. 10 days longer than females, receive 27% more food (mean meal size 396 g versus 345 g in females), grow faster, and attain higher peak and fledging masses (35). Fledging success varies between 22% and 100% (35). Chicks may become heavily infested, and probably weakened, by ticks (Ixodes auritulus) (2). Returns to colony at age 2–3 years and reaches sexual maturity at 5–6 years (2).

White-capped Albatross (steadi)

Breeding frequency and season poorly known, but recent indications are that it breeds biennially, with annual probability of breeding estimated at 0·68 (0·58–0·81). Eggs usually laid mid Nov and hatch in Feb; chicks thought to fledge in mid Aug, though a fledging period of Jun–Jul may be more likely. Permanence of pair-bonds unknown (1). Loosely colonial; nest  a mound of mud  , guano, tussock grass, herbs, feathers and other material (1). Single whitish egg  , with reddish-brown spots at large end; size similar to that of T. cauta, 74·1–118·8 mm × 63·7–74·1 mm (36); chicks  have white down  ; incubation and fledging periods unknown (1). No information concerning either juvenile survival or age of first breeding is available. Adult survival (based on mark-recapture data) estimated at 0·96 (0·91–1·00).

Conservation Status

White-capped Albatross (cauta)

Not globally threatened. Currently treated as Near Threatened. Was considered Vulnerable in Australia (37). Population decimated around turn of century by plume hunters, especially off Tasmania, where only c. 300 nests by 1909; previously c. 3300 pairs at three colonies (Albatross I, Mewstone and Pedra Branca), increasing to 5000 pairs, 7000 pairs and 200–250 pairs, respectively, in 1995 (2) and even more recently these totals were revised to 5200 pairs, 7600–12,400 pairs and 130–170 pairs, respectively, giving an overall total of 15,350 pairs and 60,000–70,000 individuals. Population on Pedra Branca may have always been small, but it appears that competition for nesting space with Australasian Gannets (Morus serrator) may steadily be reducing number of fledglings produced each year: chick production on Pedra Branca fell from over 100 to just 31 between 1993 and 2007, representing a decrease of c. 9% per year. Suffered substantial mortality at commercial fishing grounds, but benefitted from recent ban in New Zealand waters of trawlers using antiquated cable system. Thus, although ‘shy-type’ albatrosses (probably mainly T. cauta) comprised > 12% of seabirds caught by Japanese tuna longliners in Australian waters in 1989–995 (up to 900 birds per year) (38), Japanese fishing effort ceased in 1997 and current domestic effort is concentrated further N, where likelihood of encountering albatrosses is much lower. Currently, more limited overlap between distribution of adults and Australian longline fishing effort (39, 40) (total impact of trawl fisheries unknown, though could account for 75% of mortalities) (41), but juveniles from Mewstone population, at least, forage in waters off South Africa, bringing them into contact with several fisheries that pose greater threat, but no evidence that shy-type albatrosses other than steadi have been taken as bycatch in these waters (42).

White-capped Albatross (steadi)

Not globally threatened. Currently treated as Near Threatened. Annual breeding population estimated at c. 100,000 pairs in 2012 (versus estimates of 70,000–80,000 pairs in 1992–1994), with colonies on Disappointment I, Adams I and Auckland I, all in Auckland group, and Bollons I (50–100 pairs, versus just 20 pairs in mid 1990s) (43) in Antipodes group (with suspicions of breeding on Windward I) (44), with some 95% of global population breeding on Disappointment. One pair bred on Forty-Fours (Chatham Is) in Dec 1991 and Nov/Dec 1996 (36). ‘Shy-type’ albatrosses recorded in SW Atlantic for many years; most birds recorded are immature, hindering specific identification, but genetic evidence from bird on South Georgia confirmed species as T. steadi (29). Annual estimates of breeding pairs in 2007–2011 appeared to indicate very rapid population decline, but inter-annual variability and larger numbers breeding in 2012 cast some doubt on this. Need for accurate trend information highlighted by estimates of 8000 individuals taken annually as bycatch in longline and trawl fisheries, but others suggest that > 17,000 birds die per year, with c. 30% of this in New Zealand fisheries, although this percentage is apparently declining, and lack of data concerning cryptic mortality make these estimates very uncertain. Also most frequently caught species in pelagic tuna longline operations off South Africa, with an estimated 7000–11,000 killed in South African pelagic longline fishery in 1998–2000 and, in 2005, c. 500–600 ‘shy-type’ albatrosses were killed. In South African demersal trawl fishery, observer data from 2004–2005 produced an estimate of 7700 ‘shy-type’ albatrosses killed annually, and subsequent DNA analysis indicated that all were T. steadi (42). Since introduction of mandatory permit requirements in Aug 2006, whereby all vessels must deploy a bird streamer line, bycatch rate has decreased, but further data collection is required to establish new catch estimate (45). Impact of large distant-water fleets of Japan, Taiwan and Korea largely unknown, but Japanese data from 2001–2002 indicate that > 10% of recorded albatross mortalities were ‘shy-type’ (41). In Uruguayan longline fleet operating in SW Atlantic, ‘shy-type’ albatrosses comprised 25% of all birds observed in association with vessels, mostly immatures; five individuals caught as bycatch were confirmed as T. steadi, but numbers caught are not sufficient to predict an overall bycatch level for this fleet (23). Commercial exploitation of squid or fish reserves in Bass Strait could pose threat through direct competition for food. On Auckland I, nesting area was significantly reduced during 1972–1982 because of interference by pigs, and these and feral cats may also take small numbers of chicks (46).

Recommended Citation

del Hoyo, J., C. Carboneras, F. Jutglar, N. Collar, and G. M. Kirwan (2020). White-capped Albatross (Thalassarche cauta), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.whcalb1.01
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