Mount St Ysabel, Bioko.
In past, often treated as conspecific with L. fuelleborni, but the two differ in plumage (particularly juvenile), bill size and voice. Until recently considered conspecific with L. holomelas; molecular evidence#R that helps to determine separate species status for holomelas also indicates close relationship of proposed species L. willardi to present species, and willardi included here on this basis and because it shares grey irides with (at least) race camerunensis#R (eye colour of camerunensis independently confirmed in NHMUK); willardi, however, has a song consisting of four fast equal-spaced identical notes, very different from other taxa in the complex, and fuller evidence may reinstate it as a species. No information on eye colour of older proposed taxon L. melas, described from Lugege (= Nyungwe) Forest, in Rwanda, but high elevation of that forest suggests that that name is probably a junior synonym of holomelas, and does not apply to willardi. Three subspecies currently recognized.
Food and feeding
Status and conservation
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