Genetic data indicate that rufula is sister to G. blakei#R. Considerable geographical differences in vocalizations, including within nominate race, indicate that major revision of species limits is needed#R; this expected in 2017. Commonly includes G. saltuensis (which see) but this recently given species rank, confirmed here; race spatiator also once elevated to species rank#R. Six subspecies currently recognized.
Food and feeding
Status and conservation
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